Horticulture Science and Practices
Plant Anatomy, Morphology and Development
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3 TISSUE SYSTEMS OCCUR
IN PLANTS 1) Dermal Tissue System Function: Protection from the environment and water loss. Tissues: a) epidermis - single layer of cells on primary (herbaceous ) plant parts. b) periderm or bark - a corky tissue that replaces epidermis on secondary (woody) plant parts. 2) Vascular Tissue System
3) Ground or Fundamental Tissue
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1) cell wall (see next page for structure of primary and
secondary cell wall & middle lamella)
2) hemicellulose - filler between cellulose microfibrils (chains of misc. sugar) 3) pectin - cementing agent or filler; high in middle lamella and fruit; (chains of galacturonic acid); used to make jelling compounds; broken down when fruit ripens.
c) protein - mainly structural (most commonly hydroxyproline) composed of a bilayer of phospholipids and proteins; it is selectively permeable and regulates absorption into cells and leakage from cells. 3) plasmodesmata- tubular plasma membrane extensions through cell walls that connect adjacent cells. 4) cytoplasm - cytosol plus organelles; most metabolism occurs in the cytosol or its organelles.
b) organelles - specialized structures in cytoplasm, each with specific functions.
b) nucleolus - spherical, dense body; site of ribosome synthesis. 3) plastid - double membrane-bound bodies for storage and photosynthesis
2) elaioplast- fat and oil storage c) chloroplast - green plastids that contains chlorophyll*; the site of photosynthesis cytoplasm; site of protein & membrane synthesis 5) ribosome - dense spheres of RNA; protein synthesis occurs on their surface 6) vacuole- storage of organic acids, salts, anthocyanins* (blue, purple and red) pigments), metabolic wastes, enzymes and metabolites.
8) microbody- membrane-bound storage bodies with various functions. 9) microtubule - tubular rods used in mitosis & cellulose orientation in cell walls *Colored Pigments, Antioxidants and Healthy Foods Carotenoids, xanthophylls, chlorophyll & anthocyanins are antioxidants - so eat a rainbow of colors |
Structure of starch vs. cellulose
This small structural change
dramatically changes the properties
of the two compounds.Example, we can digest starch, but not cellulose (that is why we cannot eat wood!) |
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(electon micrograph (EM) of tobacco leaf cell) |
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Cell Wall Each cell is surrounded on all sides by the cell wall. The cells of herbaceous tissue (primary tissue) have only the primary cell wall and middle lamella, which are not very rigid. Most of the plant tissue we eat only has primary cell walls. When fruits ripen, the middle lamella breaks down due to pectin digestion, thus the cells slip past each other and the tissue seems softer. The cellulose microfibrils in the primary cell wall are parallel, which allows the cell wall to expand as the cell grows. The cells of woody or wood-like (lignified) tissue possess a secondary cell wall. In the secondary cell wall, the cellulose microfibrils are criss-crossed and are impregnated with lignin, both of which make secondary cell walls very hard and rigid. Generally, only xylem and sclerenchyma cells (fibers and sclereids) have secondary cell walls; all of which are dead at maturity. Cytoplasm |
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DEFINITIONS DNA (deoxyribonucleic acid)- a double helix chain of sugar-phosphates (deoxyribo sugar-phosphates) connected by nucleic acids (adenine, thymine, guanine, cytosine). RNA (ribonucleic acid) - a single stranded chain of sugar-phosphates (ribo sugar-phosphates) containing nucleic acids (adenine, uracil, guanine, cytosine). nucleic Acids - organic acids that form the base pairs of DNA and single-bases of RNA. Base Pairing of Nucleic Acids - between double strands of DNAchromosome - tightly coiled strands of DNA, which also include packing proteins to aid structure. gene - a length of DNA that codes for the production of a protein or protein subunit. - also codes for active RNAs (such as tRNA). protein - a polymer or chain of amino acids. enzyme - a protein that acts as a metabolic catalyst. Genetically Modified Organism - GMO - any organism whose genetic material has been altered using genetic engineering techniques as opposed to natural mating, mutations, recombinations or traditional plant breeding techniques. |
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HOW DO PLANTS GROW?
Plants grow from localized areas called meristems. meristem - discrete regions or groups
of cells that possess continued cell division for
the PLANTS EXHIBIT TWO TYPES OF GROWTH
a) apical meristem or apex- the growing points located at the tips of stems and roots. b)
intercalary
meristem
- the growth region at the base of grass
leaves that causes corky) tissues called the secondary plant body. Secondary growth is due to lateral meristems. lateral meristem - meristematic regions along the sides of stems and roots.
a) vascular cambium or cambium - a sheet-like meristem between the bark and wood along the sides of woody stems and roots; it gives rise to secondary xylem (commonly called wood) on the inside and secondary phloem on the outside. b)
cork
cambium or phellogen -
gives rise to the periderm
(commonly called bark). Both meristems grow away from
themselves, apical grows up in length & lateral
growth out in diameter.
Example, is a barbed wire fence nailed to a tree. |
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OF MONOCOTS, DICOTS AND GYMNOSPERMS |
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STEM MORPHOLOGY |
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bud - an underdeveloped and
unelongated stem composed of a short axis with
compressed internodes, a meristematic apex, and
primordial leaves and/or flowers. bud break - when
the bud starts to grow the bud scales peel back and the
young shoot or flowers start to emerge. axillary
bud or lateral bud - buds along side the
axis of a stem; they at were produced by the terminal
bud during growth; once they grow out and form a lateral
stem they become terminal buds of the lateral
branch. flower
bud
- a bud containing a floral meristem which develops into
flowers; usually larger than vegetative buds.
leaf
scar - a scar marking the former point of
attachment of a leaf or petiole to the stem.
internode - the part of the stem
between nodes node - part of stem marking
the point of attachment of leaves, flowers, fruits, buds
and other stems. lenticel
- rough areas on stems
(and some fruits, ex. apple) composed of loosely packed
cells extending from the cortex through the ruptured
epidermis; serve as "breathing pores" for gas exchange.
Only occur on young stems. growth rings - bud scale scars from the last terminal bud; they denote flushes of growth (usually per year). Can be used to age stems because usually 1 set of growth rings is produced per year on temperate trees in the Temperate Climatic Zone. |
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Primary Growth Vascular bundles are arranged as a ring between the cortex and pith. The pith and cortex are usually comprised of parenchyma cells. Inside each vascular bundle, the phloem is orientated towards the outside and xylem towards the inside of the stem. The outer surface is covered by the epidermis. |
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Woody Dicot or Gymnosperm Stem
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Primary Growth The vascular bundles are randomly scattered in the ground tissue (usually comprised of parenchyma cells). Each vascular bundle is surrounded by a bundle sheath and contains xylem orientated towards the inside and phloem towards the outside of the stem. The outer layer is epidermis. |
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Monocot, Dicot or Gymnosperm Root Primary Growth Root anatomy is virtually the same for monocots, dicots and gymnosperms. The vascular tissue occurs in the center, which is surrounded by two rings of cells, the pericycle and endodermis, then the cortex and epidermis. The cell walls of the endodermis that are perpendicular to the root surface (i.e. the radial and anticlinal walls) are sealed by a suberized band called the Casparian strip. Root hairs are extensions of the epidermal cells. |
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A woody dicot or gymnosperm root in secondary growth looks similar to a stem in secondary growth. Therefore, see Stem Cross Section - Woody Dicot or Gymnosperm - Secondary Growth. The main difference is there is primary xylem in the center, and the rings are not a clearly defined. |
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LEAF MORPHOLOGY |
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SIMPLE LEAF (Ficus)
tip - the terminal point of the leaf blade or lamina - the flattened,
green, margin - edge of a leaf. midrib - the most
prominent central vein in a lateral veins - secondary veins in a leaf. petiole - the leaf stalk
(connects blade to stipules - leaf-like
appendages (at the base |
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COMPOUND LEAF (rose)
leaflet - secondary leaf of a compound leaf. rachis
- an extension of the petiole bearing
petiolule - the leaflet stalk. petiole - the leaf stalk. stipules
- leaf-like appendages (at the base |
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SIMPLE LEAF - blade of the leaf
occurs as one unit (i.e. is not divided into leaflets;
however, the blade may be highly lobed or indented). |
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COMPOUND LEAF - the blade of the
leaf is divided into individual leaflets.
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Monocot Leaf (corn)
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Gymnosperm Leaf (pine
needle) |
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STOMA
stoma - an open
aperture (the stomatal pore)
in the epidermis surrounded by 2 guard cells
Mechanism of Opening
Daily Cycle Designed for gas exchange
MESOPHYLL Palisade parenchyma
Sun Grown Leaf
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FLOWER MORPHOLOGY
flower - the reproductive organ of higher plants (e.g. angiosperms or flowering plants), which contains at least 1 female reproductive part, the pistil, and/or 1 male reproductive part, the stamen.
complete- contains all floral parts, i.e. sepal, petal, stamen and pistil (Trumpet vine) incomplete - lacks one or more of the floral parts (Century oak; flowering, male flowers called catkins) perfect - contains both pistil and stamen (may or may not have sepal or petal) imperfect - lacks either pistil or stamen (may or may not have sepal or petal) pistillate (female) - contains only pistil (may or may not have sepal or petal) (cucumber) staminate (male) - contains only stamen (may or may not have sepal or petal) (cucumber) sterile - both stamen and pistil are absent, or are non-functional Plant Types Based on Flower Type
Present dioecious -
staminate (male) and pistillate (female) flowers
occur on separate plants;
Composed of 3
layers: 2) mesocarp - middle layer of the pericarp 3) endocarp - inner layer of the pericarp.
testa - protective, outer most layer of seeds; commonly called seed coat. |
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![]() (example Trumpet Vine) |
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![]() (lima bean dissection; shelled peanut) |